Email accounts are managed in the Email & Calendar tool: open it and click Email accounts in the sidebar. The same link also appears in the CRM sidebar, since one account can serve both campaign sending and the shared inbox - see The Shared Inbox and Email Accounts for how the inbox itself works.
Click New Email Account, name it, and set its from-address. Admins can mark an account organization-wide; otherwise it's personal to whoever creates it. Fill in SMTP details for sending, and IMAP details if you also want incoming mail synced - IMAP is optional and only needed to receive mail, not to send it.
Tick Campaign sending, Inbox monitoring, or both - an account can serve either purpose or both at once. An account used for inbox monitoring must be personal, since it represents one person's own mailbox.
Saving an account, or changing its host, port, credentials, or purpose, triggers a background connection test automatically. The accounts list shows the result as a status badge: Healthy, Testing while a check runs, Error with the reason if it failed, or Untested if none has run yet. Click Test next to an account any time to re-check it manually.
Campaign and inbox reply email always sends through one of your connected email accounts rather than a generic system address, so recipients see mail coming from your own domain.
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